Office Location: FSC 239
Phone Number: (717) 477-1170
Eastern New Mexico University B.S. Biology/Chemistry 1982
University of Wyoming M.S. Zoology 1985
Arizona State University Ph.D. Zoology 1995
I began teaching at Shippensburg University in 1996. Prior
to that, I moved around the western United States, earning my BS, MS, and PhD
in New Mexico, Wyoming, and Arizona respectively. In between my MS and PhD
studies, I spent six years teaching junior high and high school science and
coaching track and cross country in New Mexico. Although I am interested in
anything dealing with ecology or evolution, my real fascination is with the
ecology of salamanders, turtles, and vernal ponds. I also have a strong
interest in conservation biology, as human impacts threaten the future of the
organisms and wild places that I cherish. My wife and I live on a small farm
north of Shippensburg, where I spend my much of my spare time enjoying the
natural world and restoring native habitats.
Ecology and evolution
Conservation biology of amphibians and turtles
Ecology of vernal ponds
Principles of Evolution
Current Graduate Students
Kelsey Miller – The role of red-spotted newts, (Notophthalmus viridescens), in aquatic
Sarah Kropf – The effects of the neonicotinoid
insecticide Clothianidin on survival, growth, and development of larval
amphibians and aquatic food webs.
Paul Mashett – The effects of tannins on predator
detection in larval amphibians and predator-prey interactions in aquatic food
Alex Shupinski – The effects of grazing within
wetland easements on vegetation structure and occupancy, abundance and species
richness of birds.
Hanna Roos - The effects of grazing within wetland
easements on habitat structure and habitat use and movements of turtles.
Recent Undergraduate Student Research Projects
Kate Destafano, Alyssa Hay, and
Emily Sarver, 2014-2017 - Effect of an invasive plant species, Ficaria verna, on a native woodland
species, Mertensia virginica, in
floodplain forest in south Central Pennsylvania.
Brandon Mullen and Drew Yauneridge,
2015-2017 - Effects of forest change on leaf processing in vernal pond ecosystems
Sam Gowan, 2016-2017 - Analysis of
the ecology and diet of eastern newts in vernal pond communities in south
Emily Sarver, Paul Christman, Brandon
Mullen, Riley Papson, and Alyssa Hay, 2015-2016 -Ecological study of amphibian
and reptile populations in a restored wetland in southcentral Pennsylvania.
Nathan Alexander, 2013 - Population
monitoring of spotted turtles, Clemmys
guttata, a species of conservation concern in Pennsylvania
Student Research Projects
Karen Schwander, 2015-2016. The
Effect of Native and Exotic Invasive Plants on Vernal Pool Composition and
Amphibian Growth and Survival
Miranda McCleaf, 2014-2015. An
Assessment of the Movement and Habitat Use of Spotted Turtles (Clemmys guttata)
during Regrowth of a Managed Area in Pennsylvania
Katie Seymore, 2013-2014. The effect
of forest composition and caddisfly larvae (Limnephilus
indivisus) on vernal pond communities.
Justin Atkins, 2009-2010. The
effects of hypoxia on interactions between an aquatic predator (Notonecta
irrorata) and its amphibian prey (Ambystoma maculatum and Lithobates
Amy Williams, 2006-2007 - The effect
of increased salinity from road de-icing salts on predatory interactions among
three amphibian species.
Mehring, A. S.*
and T. J. Maret. 2011. Red maple dominance enhances fungal and shredder growth
and litter processing in temporary ponds. Limnology and Oceanography
Mott, C. L.*
and T. J. Maret. 2011. Species-specific Patterns of Agonistic Behavior among
Larvae of Three Syntopic Species of Ambystomatid Salamanders. Copeia 2011:9-17.
Steele, M. A.,
M. C. Brittingham, T. J. Maret, and J. F. Merritt, eds. 2010. Terrestrial
Vertebrates of Pennsylvania: A Complete Guide to Species of Conservation
Concern. Johns Hopkins University Press, Baltimore, Maryland.
Morris, K. M.*
and T. J. Maret. 2007. Effects of timber management on pond breeding
salamanders. Journal of Wildlife Management 71:1034-1041.
Maret, T. J.,
J. D. Snyder, and J. P. Collins. 2006. Altered drying regime controls
distribution of endangered salamanders and introduced predators. Biological
* graduate student