/uploadedImages/Ship/Biology/Dr_Maret.jpgTim Maret


Office Location: FSC 239

Phone Number: (717) 477-1170

Email: tjmare@ship.edu


Eastern New Mexico University B.S. Biology/Chemistry 1982

University of Wyoming M.S. Zoology 1985

Arizona State University Ph.D. Zoology 1995


I began teaching at Shippensburg University in 1996. Prior to that, I moved around the western United States, earning my BS, MS, and PhD in New Mexico, Wyoming, and Arizona respectively. In between my MS and PhD studies, I spent six years teaching junior high and high school science and coaching track and cross country in New Mexico. Although I am interested in anything dealing with ecology or evolution, my real fascination is with the ecology of salamanders, turtles, and vernal ponds. I also have a strong interest in conservation biology, as human impacts threaten the future of the organisms and wild places that I cherish. My wife and I live on a small farm north of Shippensburg, where I spend my much of my spare time enjoying the natural world and restoring native habitats.

Research Interests

Ecology and evolution

Conservation biology of amphibians and turtles

Ecology of vernal ponds

Courses Taught


Principles of Evolution

Conservation Biology

Environmental Biology

Current Graduate Students

Kelsey Miller – The role of red-spotted newts, (Notophthalmus viridescens), in aquatic food webs.


Sarah Kropf – The effects of the neonicotinoid insecticide Clothianidin on survival, growth, and development of larval amphibians and aquatic food webs.


Paul Mashett – The effects of tannins on predator detection in larval amphibians and predator-prey interactions in aquatic food webs.


Alex Shupinski – The effects of grazing within wetland easements on vegetation structure and occupancy, abundance and species richness of birds.


Hanna Roos - The effects of grazing within wetland easements on habitat structure and habitat use and movements of turtles.

Recent Undergraduate Student Research Projects


Kate Destafano, Alyssa Hay, and Emily Sarver, 2014-2017 - Effect of an invasive plant species, Ficaria verna, on a native woodland species, Mertensia virginica, in floodplain forest in south Central Pennsylvania.


Brandon Mullen and Drew Yauneridge, 2015-2017 - Effects of forest change on leaf processing in vernal pond ecosystems


Sam Gowan, 2016-2017 - Analysis of the ecology and diet of eastern newts in vernal pond communities in south central Pennsylvania


Emily Sarver, Paul Christman, Brandon Mullen, Riley Papson, and Alyssa Hay, 2015-2016 -Ecological study of amphibian and reptile populations in a restored wetland in southcentral Pennsylvania.


Nathan Alexander, 2013 - Population monitoring of spotted turtles, Clemmys guttata, a species of conservation concern in Pennsylvania 

Recent Graduate Student Research Projects

Karen Schwander, 2015-2016. The Effect of Native and Exotic Invasive Plants on Vernal Pool Composition and Amphibian Growth and Survival


Miranda McCleaf, 2014-2015. An Assessment of the Movement and Habitat Use of Spotted Turtles (Clemmys guttata) during Regrowth of a Managed Area in Pennsylvania


Katie Seymore, 2013-2014. The effect of forest composition and caddisfly larvae (Limnephilus indivisus) on vernal pond communities.


Justin Atkins, 2009-2010. The effects of hypoxia on interactions between an aquatic predator (Notonecta irrorata) and its amphibian prey (Ambystoma maculatum and Lithobates sylvaticus).


Amy Williams, 2006-2007 - The effect of increased salinity from road de-icing salts on predatory interactions among three amphibian species.


Selected Recent Publications

Mehring, A. S.* and T. J. Maret. 2011. Red maple dominance enhances fungal and shredder growth and litter processing in temporary ponds. Limnology and Oceanography 56:1106–1114.

Mott, C. L.* and T. J. Maret. 2011. Species-specific Patterns of Agonistic Behavior among Larvae of Three Syntopic Species of Ambystomatid Salamanders. Copeia 2011:9-17.

Steele, M. A., M. C. Brittingham, T. J. Maret, and J. F. Merritt, eds. 2010. Terrestrial Vertebrates of Pennsylvania: A Complete Guide to Species of Conservation Concern. Johns Hopkins University Press, Baltimore, Maryland.

Morris, K. M.* and T. J. Maret. 2007. Effects of timber management on pond breeding salamanders. Journal of Wildlife Management 71:1034-1041.

Maret, T. J., J. D. Snyder, and J. P. Collins. 2006. Altered drying regime controls distribution of endangered salamanders and introduced predators. Biological Conservation 127:129-138.

* graduate student