/uploadedImages/Ship/Biology/Dr_Maret.jpgTim Maret


Office Location: FSC 239

Phone Number: (717) 477-1170

Email: tjmare@ship.edu

Personal/Research Webpage: http://webspace.ship.edu/tjmare/


Eastern New Mexico University B.S. Biology/Chemistry 1982

University of Wyoming M.S. Zoology 1985

Arizona State University Ph.D. Zoology 1995


I began teaching at Shippensburg University in 1996. Prior to that, I moved around the western United States, earning my BS, MS, and PhD in New Mexico, Wyoming, and Arizona respectively. In between my MS and PhD studies, I spent six years teaching junior high and high school science and coaching track and cross country in New Mexico. Although I am interested in anything dealing with ecology or evolution, my real fascination is with the ecology of salamanders, turtles, and vernal ponds. I also have a strong interest in conservation biology, as human impacts threaten the future of the organisms and wild places that I cherish. My wife and I live on a small farm north of Shippensburg, where I spend my much of my spare time enjoying the natural world and restoring native habitats.

Research Interests

Ecology and evolution

Conservation biology of amphibians and turtles

Ecology of vernal ponds

Roles of competition and predation in regulating populations

Courses Taught


Principles of Evolution

Conservation Biology

Environmental Biology

Recent Undergraduate Student Research Projects

Ben Ahlswede and Stephanie Summers, 2012 - Interspecific interactions and terrestrial ecology of adult Ambystomatid salamanders.

Katie Weaver and Evan Smith, 2012 - Developing an in-house PCR to test for the presence of Batrachochytrium dendrobatidis (with Drs. Lehman and Bumbaugh).

Katie Weaver, James Francis, Lawrence Ashton, Annie Birdsong, Hayley Crowell, Grant Izzo, and Justin Wissler, 2010-2011 - Survey for the presence and prevalence of Batrachochytrium dendrobatidis in south central Pennsylvania

Lawrence Ashton and Grant Izzo, 2009 - The Effects of timber harvest on small mammals in Michaux State Forest, Pennsylvania.

Danille Dougherty, 2007 - The effects of the insecticide methoprene on Jefferson salamander and wood frog larvae.

Recent MS Student Research Projects

Justin Atkins, 2009-2010 - The effects of hypoxia on interactions between an aquatic predator (Notonecta irrorata) and its amphibian prey (Ambystoma maculatum and Lithobates sylvaticus).

Amy Williams, 2006-2007 - The effect of increased salinity from road de-icing salts on predatory interactions among three amphibian species.

John Chripczuk, 2005-2006 - Effects of Notonecta (Hemiptera: Notonectidae) on amphibian assemblages in vernal ponds.

James Mellon, 2005-2006 - Hatchling ecology and nest predation in the northern diamond terrapin, Malaclemys terrapin terrapin.

Jason Ambler (with Dr. Lehman), 2003-2006 - Determination of population structure of three vernal pond amphibians using molecular techniques.

Selected Recent Publications

Mehring, A. S.* and T. J. Maret. 2011. Red maple dominance enhances fungal and shredder growth and litter processing in temporary ponds. Limnology and Oceanography 56:1106–1114.

Mott, C. L.* and T. J. Maret. 2011. Species-specific Patterns of Agonistic Behavior among Larvae of Three Syntopic Species of Ambystomatid Salamanders. Copeia 2011:9-17.

Steele, M. A., M. C. Brittingham, T. J. Maret, and J. F. Merritt, eds. 2010. Terrestrial Vertebrates of Pennsylvania: A Complete Guide to Species of Conservation Concern. Johns Hopkins University Press, Baltimore, Maryland.

Morris, K. M.* and T. J. Maret. 2007. Effects of timber management on pond breeding salamanders. Journal of Wildlife Management 71:1034-1041.

Maret, T. J., J. D. Snyder, and J. P. Collins. 2006. Altered drying regime controls distribution of endangered salamanders and introduced predators. Biological Conservation 127:129-138.

* graduate student