/uploadedImages/Ship/Biology/Dr_Maret.jpgTim Maret


Office Location: FSC 239

Phone Number: (717) 477-1170

Email: tjmare@ship.edu


Eastern New Mexico University B.S. Biology/Chemistry 1982

University of Wyoming M.S. Zoology 1985

Arizona State University Ph.D. Zoology 1995


I began teaching at Shippensburg University in 1996. Prior to that, I moved around the western United States, earning my BS, MS, and PhD in New Mexico, Wyoming, and Arizona respectively. In between my MS and PhD studies, I spent six years teaching junior high and high school science and coaching track and cross country in northern New Mexico. Although I am interested in anything dealing with ecology or evolution, my real fascination is with the ecology of salamanders, turtles, and vernal ponds. I also have a strong interest in conservation biology, as human impacts threaten the future of the organisms and wild places that I cherish. My wife and I live on a small farm north of Shippensburg, where I spend much of my spare time enjoying the natural world and restoring native habitats.

Research Interests

Ecology and evolution

Conservation biology of amphibians and turtles

Ecology of vernal ponds

Courses Taught

BIO 242 Ecology

BIO 430 Principles of Evolution

BIO 444 Conservation Biology

BIO 522 Community Ecology

Recent Student Research

Alex Shupinski 2017-2018 – The effects of grazing within wetland easements on vegetation structure and occupancy, abundance and species richness of birds.

Hanna Roos 2017-2018 - Habitat use, movement, and home range of bog turtles (Glyptemys muhlenbergii) in southeastern, PA

Claudia Knutelski 2017-2018 - The impact of population density on survivorship and metamorphosis in wood frogs (Lithobates sylvaticus).

Sam Govan 2016-2018 - Factors influencing the diet of the red-spotted newt, Notophthalmus viridescens, in vernal pools in southcentral Pennsylvania

Joshua Frost 2017-2018 - Examining pollinator diversity in cattle-grazed and ungrazed wetlands.

Kate Destafano, Alyssa Hay, Emily Sarver, Patrick Boddicker 2014-2018 - Effect of an invasive plant species, Ficaria verna, on a native woodland species, Mertensia virginica, in floodplain forest in south Central Pennsylvania.

Sarah Kropf 2016-2017 - The effects of the neonicotinoid insecticide Clothianidin on survival, growth, and development of larval amphibians and aquatic food webs.

Paul Mashett 2016-2017 – The effects of tannins on predator detection in larval amphibians and predator-prey interactions in aquatic food webs.

Brandon Mullen 2015-2017 - Effects of forest change on leaf processing in vernal pond ecosystems

Karen Schwander, 2015-2016. The Effect of Native and Exotic Invasive Plants on Vernal Pool Composition and Amphibian Growth and Survival

Miranda McCleaf, 2014-2015. An Assessment of the Movement and Habitat Use of Spotted Turtles (Clemmys guttata) during Regrowth of a Managed Area in Pennsylvania

Katie Seymore, 2013-2014. The effect of forest composition and caddisfly larvae (Limnephilus indivisus) on vernal pond communities.

Selected Publications

Mehring, A. S.* and T. J. Maret. 2011. Red maple dominance enhances fungal and shredder growth and litter processing in temporary ponds. Limnology and Oceanography 56:1106–1114.

Mott, C. L.* and T. J. Maret. 2011. Species-specific Patterns of Agonistic Behavior among Larvae of Three Syntopic Species of Ambystomatid Salamanders. Copeia 2011:9-17.

Steele, M. A., M. C. Brittingham, T. J. Maret, and J. F. Merritt, eds. 2010. Terrestrial Vertebrates of Pennsylvania: A Complete Guide to Species of Conservation Concern. Johns Hopkins University Press, Baltimore, Maryland.

Morris, K. M.* and T. J. Maret. 2007. Effects of timber management on pond breeding salamanders. Journal of Wildlife Management 71:1034-1041.

Maret, T. J., J. D. Snyder, and J. P. Collins. 2006. Altered drying regime controls distribution of endangered salamanders and introduced predators. Biological Conservation 127:129-138.

Maret, T. J., and J. P. Collins. 1997. Ecological origin of morphological diversity: a study of alternative trophic phenotypes in larval salamanders. Evolution 5:898-905.

Maret T. J. and J. P. Collins. 1996. Effect of prey vulnerability on population size structure of a gape-limited predator. Ecology 77:320-324.

* student